Acalyptris pistaciae

Difference from A. minimella:

• Absence of a central yellow spot on the forewing underside and absence of costal bristles;
• Hairpencil more distinct in A. pistaciae
• Male genitalia characterized by medial toothed lobe of valva in contrast to the basal lobe in minimella, and by the U-shaped ventral process;
• Female genitalia are distinguished by the elaborate vaginal sclerotisation with three processes and the 21⁄2 convolutions in the ductus spermathecae (3 in minimella).

Male Habitus:

• Forewing length 1.8–2.4 mm, wingspan 4.0–5.3 mm.
• Frontal tuft ochreous white to yellowish white, occasionally with few fuscous scales.
• Scape white, antenna with 27–33 segments
• Thorax and forewing ochreous, irrorate with few light brown scales, cilia white, cilia line indistinct; underside without yellow spot; basalmost dorsal cilia forming long hairpencil.
• Hindwing grey, costal bristles absent; distinct, yellowish brown hairpencil arising near frenulum; mixing with forewing hairpencil; underside yellow basally. Abdomen yellow with darker middorsal line; anal tufts grey.

Female Habitus:

• Forewing length 1.6–2.2 mm, wingspan 3.7–4.9 mm.
• Antenna with 20–30 segments.
• Hairpencils absent.
• Abdomen dorsally brown, abdominal tip broadly rounded.

Male Genitalia:

• Vinculum anteriorly concave, ventral plate not large.
• Tegumen small, triangular; uncus band-shaped, with indistinct central process, in lateral view split distally; gnathos with long narrow central element.
• Valva narrow, with medial inner lobe, with variably toothed margin and a smaller lobe dorsally, hidden by ventral lobe.
• Transtilla without transverse bar, sublateral process distinct.
• Aedeagus with bifurcate ventral carina or with 4–5 lobes, ventrally tightly fused to U-shaped ventral process; a pair of curved dorsal carinae present; vesica with large curved cornutus and basal sclerotisation associated with cathrema; many smaller cornuti.

Female Genitalia:

• T8 narrow, with produced lateral corners, two small groups of scales and some setae.
• Anal papillae conspicuous, with 15–20 setae; apophyses straight, of approximately same length.
• Total length of bursa ca. 530–760 μ m.
• Vestibulum with elaborate sclerotisations, more or less forming ring and three apophyses, partly serrate.
• Ductus bursae with many pectinations; corpus bursae without pectinations; with reticulate signa, shortest ca. 285–325 μ m, longest 310–420 μ m long, margin crenate and narrow, 2–3 cells wide.
• Ductus spermathecae with 21⁄2 convolutions.

Larva (final instar):

• Head capsule ca. 315–380 μ m wide.
• Mandibular cusps blunt.
• A9 with 3 pairs of setae.
• Integument covered with extremely short microtrichia (ca 1 μ m long) on all segments.
• No obvious differences with A. minimella.