Acalyptris platani

Male Habitus:

• Males unmistakeable by colour pattern and cuspidate hindwings with special raised scales, not resembling any other Nepticulidae.
• Forewing length 2.4–2.5 mm, wingspan 5.2–5.4 mm.
• Frontal tuft yellowish, on vertex fuscous, two areas clearly separated; collar very inconspicuous, as vertex; scape and pedicel white.
• Antenna with ca. 34 segments.
• Thorax yellowish white to brown.
• Forewing basal 1/4 yellowish brown, followed by broad yellowish white fascia to 1/2; distal part of wing brown, yellowish white opposite tornal and costal spots, sometimes united in a second fascia; cilia silvery white, cilia line distinct.
• Underside forewing basal 2/3 (in rest in contact with hindwing androconial patch) pale, with very few elongate brown scales only, distal third brown.
• Hindwing very broad at base, clearly cuspidate towards tip; costal bristles present; upper surface of basal 2/3 covered with a thick indumentum of white raised special scales, interrupted along midline by a distinct, straight, longitudinal furrow, devoid of scales; costa with a row of short yellowish brown hair scales instead of cilia.
• Underside white.
• Abdomen with yellow inconspicuous anal tufts.

Male Genitalia:

• Male genitalia very characteristic, separated from the other species in the group by the valva with two inner lobes.
• Vinculum anteriorly and posteriorly concave.
• Tegumen rounded, forming obtuse pseuduncus; uncus band-shaped, with indistinct central process, with
  pair of setae; gnathos with long, pointed central element.
• Valva at base with long inner process, subterminally with inner process on dorsal surface, apex rounded.
• Transtilla without transverse bar.
• Aedeagus with ventral carinae ending in forked lobes, tightly fused to ventral process; pair of lateral, pointed carinae present; vesica with numerous small cornuti and one large, serrate cornutus.

Female Habitus:

• Forewing length 2.3–2.4 mm, wingspan 5.0–5.4 mm.
• Antenna with 27–29 segments.
• Forewing underside pale grey-brown, normally covered with scales.
• Hindwing normal, without special scales.

Female Genitalia:

• Female genitalia with vaginal sclerotisations which are characteristic for the group, separated from the next three species by the more rounded abdominal tip and extremely narrow signa with crenate margins.
• T8 with few scales and setae only.
• Anal papillae with 19–21 setae.
• Vestibulum with elaborate sclerotisations, partly serrate and densely covered with pectinations.
• Total bursa length ca. 765 μm.
• Corpus bursae elongate, without pectinations, with very narrow and long reticulate signa (length ca. 420–450 μ m), margins crenate.
• Ductus spermathecae with 2 convolutions and long and conspicuous vesicle.

Larva (final)

• Head capsule ca. 355–390 μ m wide.
• Mandibular cusps blunt.
• A9 with 2 pairs of setae.
• Integument covered with extremely short microtrichia (ca 1 μ m long), absent on A9 and 10. 
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